2Dept of Pulmonary Diseases, University Medical Centre Groningen, GRIAC (Groningen Research Institute for Asthma and COPD), University of Groningen, Groningen, The Netherlands
3Dept of Pathology and Medical Biology, University Medical Centre Groningen, GRIAC, University of Groningen, Groningen, The Netherlands
4Woolcock Institute of Medical Research, The University of Sydney, Glebe, NSW, Australia
5School of Life Sciences, University of Technology, Sydney, NSW, Australia